下述哪些疾病可引起肝细胞性黄疸（）

A.钩端螺旋体病

B.病毒性肝炎

C.伤寒，副伤寒

D.革兰阴性杆菌败血症

E.细菌性痢疾

To keep healthy,Professor Johnson           cycling as a regular form of exercise after he retired.

A. took up

B. caught on    

C. carried out

D. made for